Key words: density, flight height, occurrence patterns, seabirds, offshore wind energy
References
Adams, J., Felis, J., Mason, J. W., & Takekawa, J. Y. (2014).
Pacific continental shelf environmental assessment (PaCSEA): Aerial seabird and marine mammal surveys off northern California, Oregon, and Washington, 2011-2012 (OCS Study BOEM 2014-003).
http://www.boem.gov/2014-003/
Ainley, D. G. (1976). The occurrence of seabirds in the coastal region of California. Western Birds, 7(2), 33-68.
Ainley, D. G., & Boekelheide, R. J. (Eds.). (1990). Seabirds of the Farallon Islands: Ecology, structure and dynamics of an upwelling system community. Stanford University Press.
Ainley, D. G., & Hyrenbach, K. D. (2010). Top-down and bottom-up factors affecting seabird population trends in the California current system (1985-2006).
Progress in Oceanography,
84(3-4), 242-254.
https://doi.org/10.1016/j.pocean.2009.10.001
Ainley, D. G., Dugger, K. D., Ford, R. G., Pierce, S. D., Reese, D. C., Brodeur, R. D., Tynan, C. T., & Barth, J. A. (2009). The spatial association of predators and prey at frontal features in the northern California Current: Competition, facilitation, or merely co-occurrence?
Marine Ecology Progress Series,
389, 271-294.
https://doi.org/10.3354/meps08125
Ainley, D. G., McIver, W., Adams, J., & Parker, M. (2020). Ashy Storm-Petrel (
Oceanodroma homochroa). In P. G. Rodewald (Ed.),
Birds of the world (version 1.0.). Cornell Lab of Ornithology.
https://doi.org/10.2173/bow.asspet.01
Ainley, D. G., Nettleship, D. N., & Storey, A. E. (2021). Common Murre (
Uria aalge). In S. M. Billerman, P. G. Rodewald, & B. K. Keeney (Eds.),
Birds of the world (version 2.0). Cornell Lab of Ornithology.
https://doi.org/10.2173/bow.commur.02
Ainley, D. G., Porzig, E., Zajanc, D., & Spear, L. B. (2015). Seabird flight behavior and height in response to altered wind strength and direction.
Marine Ornithology,
43(1), 25-36.
https://doi.org/10.5038/2074-1235.43.1.1098
Ainley, D. G., Spear, L. B., Tynan, C. T., Barth, J. A., Pierce, S. D., Ford, G. R., & Cowles T. J. (2005). Physical and biological variables affecting seabird distributions during the upwelling season of the northern California Current.
Deep-Sea Research II,
52(1-2), 123-143.
https://doi.org/10.1016/j.dsr2.2004.08.016
Bates, D., Machler, M., Bolker, B., & Walker, S. (2015). Fitting linear mixed-effects models using lme4.
Journal of Statistical Software,
67(1), 1-48.
https://doi.org/10.18637/jss.v067.i01
Bograd, S. J., Schroeder, I., Sarkar, N., Qiu, X., Sydeman, W. J., & Schwing F. B. (2009). Phenology of coastal upwelling in the California Current.
Geophysical Research Letters, 36, 1-5.
https://doi.org/10.1029/2008GL035933
Bolin, R. L., & Abbott, D. P. (1963). Studies on the marine climate and phytoplankton of the coast of California 1954-1960. California Cooperative Oceanic Fisheries Investigations Reports, 9, 23-45.
Briggs, K. T., Tyler, W. B., Lewis, D. B., & Carlson, D. R. (1987). Bird communities at sea off California: 1975 to 1983.
Studies in Avian Biology,
11, 1-74.
https://doi.org/10.2307/4087784
Carboneras, C., Jutglar, F., & Kirwan, G. M. (2020). Short-tailed Albatross (
Phoebastria albatrus). In J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, & E. de Juana (Eds.),
Birds of the world (version 1.0.). Cornell Lab of Ornithology.
https://doi.org/10.2173/bow.shtalb.01
Chelton, D. B., Bernal, P. A., & McGowan, J. A. (1982). Large-scale interannual physical and biological interaction in the California Current. Journal of Marine Research, 40(4), 1095-1125.
Clarke, E. D., Spear, L. B. McCracken, M. L., Marques, F. F. C., Borchers, D. L., Buckland, S. T., & Ainley, D. G. (2003). Validating the use of generalized additive models and at-sea surveys to estimate size and temporal trends of seabird populations.
Journal of Applied Ecology,
40(2), 278-292.
https://doi.org/10.1046/j.1365-2664.2003.00802.x
Cook, A. S. C. P., Ward, R. M., Hansen, W. S., & Larsen, L. (2018).
Estimating seabird flight height using LiDAR. (Scottish Marine and Freshwater Science, 9(14))
https://doi.org/10.7489/12131-1
Ford, R. G., Terrill, S., Casey, J., Shearwater, D., Schneider, S. R., Ballance, L. T., Terrill, L., Tollefson, M., & Ainley, D. G. (2021). Distribution patterns and population size of the ashy storm petrel Oceanodroma homochroa. Marine Ornithology, 49(2), 193-204. http://doi.org/10.5038/2074-1235.49.2.1424
Grémillet, D., Ponchon, A., Paleczny, M., Palomares, M-L. D., Karpouzi, V., & Pauly, D. (2018). Persisting worldwide seabird-fishery competition despite seabird community decline.
Current Biology,
28(24), 4009-4013.
https://doi.org/10.1016/j.cub.2018.10.051
Harwood, A. J., Perrow, M. R., & Berridge, R. J. (2018). Use of an optical rangefinder to assess the reliability of seabird flight heights from boat-based surveyors: Implications for collision risk at offshore wind farms. Journal of Field Ornithology, 89(4), 372-383. https://doi.org/10.1111/jofo.12269
Joyce, T. W. (2016).
Foraging ecology, biogeography, and population biology of seabird and toothed whale predators in the Anthropocene. [Doctoral dissertation, University of California San Diego].
https://escholarship.org/uc/item/0zp36438
Kelsey, E. C., Felis, J. J., Pereksta, D. M., & Adams, J. (2025).
Revised marine bird collision and displacement vulnerability index for U.S. Pacific Outer Continental Shelf offshore wind energy development (Data Report 1214). Bureau of Ocean Energy Management.
https://doi.org/10.3133/dr1214
Largey, N., Cook, A. S. C. P., Thaxter, C. B., McCluskie, A., Stokke, B. G., Wilson, B., & Masden, E. A. (2021). Methods to quantify avian airspace use in relation to wind energy development.
Ibis,
163(3), 747-764.
https://doi.org/10.1111/ibi.12913
Leirness, J. B., Adams, J., Ballance, L. T., Coyne, M., Felis, J. J., Joyce, T., Pereksta, D. M., Winship, A. J., Jeffrey, C. F. G., Ainley, D., Croll, D., Evenson, J., Jahncke, J., McIver, W., Miller, P. I., Pearson, S., Strong, C., Sydeman, W., Waddell, J. E., … Christensen, J. (2021).
Modeling at-sea density of marine birds to support renewable energy planning on the Pacific Outer Continental Shelf of the contiguous United States (OCS Study BOEM 2021-014). Bureau of Ocean Energy Management.
https://espis.boem.gov/final%20reports/BOEM_2021-014.pdf
Li, B., Lingsma, H. F., Steyerberg, E. W., & Lesaffre, E. (2011). Logistic random effects regression models: A comparison of statistical packages for binary and ordinal outcomes.
BMC Medical Research Methodology,
11(77), 1-11.
https://doi.org/10.1186/1471-2288-11-77
Mason, J. W., Mcchesney, G. J., McIver, W. R., Carter, H. R., Takekawa, J. Y., Golightly, R. T., Ackerman, J. T., Orthmeyer, D. L., Perry, W. M., Yee, J. L., Pierson, M. O., & McCrary, M. D. (2007). At-sea distribution and abundance of seabirds off southern California: A 20-year comparison. Studies in Avian Biology, 33, 1-101.
Nur, N., Jahncke, J., Herzog, M. P., Howar, J., Hyrenbach, K. D., Zamon, J. E., Ainley, D. G., Wiens, J. A., Morgan, K., Ballance, L. T., & Stralberg, D. (2011). Where the wild things are: Predicting hotspots of seabird aggregations in the California Current System.
Ecological Applications,
21(6), 2241-2257.
https://doi.org/10.1890/10-1460.1
Paleczny, M., Hammill, E., Karpouzi, V., & Pauly, D. (2015). Population trend of the world's monitored seabirds, 1950-2010.
PLOS One,
10(6), Article e0129342.
https://doi.org/10.1371/journal.pone.0129342
Pennycuick, C. J. (1987a). Flight of seabirds. In J. P. Croxall (Ed.), Seabirds: Feeding biology and role in marine ecosystems (pp. 43-62). Cambridge University Press.
Pennycuick, C. J. (1987b). Flight of auks (alcidae) and other northern seabirds compared with southern procellariiformes: Ornithodolite observations.
Journal of Experimental Biology,
128(1), 335-347.
https://doi.org/10.1242/jeb.128.1.335
R Core Team. (2016).
R: A language and environment for statistical computing. R Foundation for Statistical Computing.
https://www.R-project.org/
Russell, T. M., Szesciorka, A. R., Joyce, T., Ainley, D. G., & Ballance, L. T. (2023). National Marine Sanctuaries capture enhanced abundance and diversity of the California Current Ecosystem avifauna.
Journal of Marine Systems,
240(1038887), 1-16.
https://doi.org/10.1016/j.jmarsys.2023.103887
Schneider, S. R., Kramer, S. H., Bernstein, S. B., Terrill, S. B., Ainley, D. G., & Matzner, S. (2024) Autonomous thermal tracking reveals spatiotemporal patterns of seabird activity relevant to interactions with floating offshore wind facilities.
Frontiers in Marine Science,
11, 1346758.
https://doi.org/10.3389/fmars.2024.1346758
Severy, M., Ortega, C., Chamberlin, C., & Jacobson, A. (2020).
Wind speed resource and power generation profile (California North Coast Offshore Wind Studies). Schatz Energy Research Center.
https://schatzcenter.org/pubs/2020-OSW-R2.pdf
Simons, T. R., & Bailey, C. N. (2020). Hawaiian Petrel (
Pterodroma sandwichensis). In A. F. Poole, & F. B. Gili (Eds.),
Birds of the world (version 1.0.). Cornell Lab of Ornithology.
https://doi.org/10.2173/bow.hawpet1.01.1
Spear, L. B., Ainley, D. G., Howell, S. N. G., Hardesty, B. D., & Webb, S. G. (2004). Reducing biases affecting at-sea surveys of seabirds: Use of multiple observer teams. Marine Ornithology, 32(2), 147-157. https://doi.org/10.5038/2074-1235.32.2.616
Spear, L. B., Nur, N., & Ainley, D. G. (1992). Estimating absolute densities of flying seabirds using analyses of relative movement. The Auk, 109(2), 385-389.
Veit, R. R., McGowan, J. A., Ainley, D. G., Wahl, T. R., & Pyle, P. (1997). Apex marine predator declines 90% in association with changing oceanic climate.
Global Change Biology,
3(1), 23-28.
https://doi.org/10.1046/j.1365-2486.1997.d01-130.x
Webb, A., & Nehls, G. (2019). Surveying seabirds. In D. M. Pereksta (Ed.),
Wildlife and wind farms, conflicts and solutions (pp. 60-96). Pelagic Publishing.
https://doi.org/10.2307/jj.29010223.7